i ,'.' catálogo de germoplasma de frijol común phaseolus...
TRANSCRIPT
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il)!flli;j¡, I:!Sf()H(h ¡ Catálogo --------------- de Germoplasma
Unidad
de Frijol común Phaseolus vulgaris L.
de Recursos Genéticos Cali - Colombia Nov. 1984
El ClAT es una institución sin ánimo de lucro, dedicada al desaITOllo agricola y económico de las zonas tropicales bajas. Su sede principal se encuentra en un terreno de 522 hectáreas" cercano a Cali, Colombia. Dícho terreno es propiedad del gobierno colombiano, el cual, en su calidad de anfitrión, brinda apoyo a las actívídades del elATo Este dispone, igualmente, de dos subestaciones propiedad de la Fundación para la Educación SuperioríFESI: QuHichao, con una extensión de 184 hectáreas, y Popayán, con 73 hectáreas, y de una subestación de 30 hectáreas-CIAT -Santa Rosa~ ubicada en terrenos cedidos por la FederaCÍón de Arroceros de Colombia IFEDEARROZl, cerca a ViHavicencio. Junto con el Instituto Colombiano Agropecuario IICAI, el CIAT administra el Centro Nacional de Investigaciones Agropecuarias Carimagua, de 22,000 hectáreas, en los Llanos Orientales y colabora con el mismo JeA en varias de sus otras estaciones experimentales en Colombia. El CIAT también lleva a cabo investigaciones en varias sedes de instituciones agrícolas nacionales en otros países de América Latina. Los programas del CIAT son financiados por un grupo de donantes que en su mayoría pertenecen al Grupo Consultivo para la Investigación Agrícola Internacional (CGIAR). Durante 1984 tales donantes son los gobiernos de Australia, Bélgica, Canadá, España, Estados Unidos de América, Francia, Holanda, Italia, Japón, Noru~ega, el Reino Unido) la República Federal de Alemania, Suecia y S4izaj la Agencia Alemana de Cooperación Técnica IGTZ); el .Banco Interamericano ·de De· sarrollo (BID); el Banco Mundial; el Centro Internacional de Investigaciones para el DesaITOUo ICIlDI; la Comunidad Económica Europea ¡CEE); el Fondo Internacional para el Desarrollo Agrícola (lFAD); el·Fondo de la OPEP para el OesaITOUo Internacional; la Fundación Ford; la Fundación Rockefeller; la Fundación W. K. Kellogg; la Organización de las Naciones Unidas para la Agricultura y la Alimentación (F AO), y el Programa de las Naciones Vnidas para el Desarrollo IPNUDJ.
La infonnación y las conclusiones contenidas en esta publicación no reflejan, necesariamente, el punto de vista de las entidades mencionadas anterionnente.
. .'
Catálogo
I SN N 0120-6265 Nov. 1984
de germoplasma de frijol común . Phaseolus vulgaris L.
"'~" ~~-; ;;. ;, , ~--:-..
\ i r
Editor Rigoberto Hidalgo, MS
cc~~u Centro Internacional de Agricultura Tropical
Colaboradores
Caracterización. Evaluación: Introducción germoplasma: Procesamiento datos: Conservación: Analista de Sistemas:
Hember Rubiano, Ing. Agr. Orlando Toro, Tec. Agr. }ulieta Ortiz Daniel Morales María del Rosario Henao, Ing. S.e.
Centro Internacional de Agricultura Tropical ClAT Apartado Aéreo 6713 Cali - Colombia 300 copias Impreso en Colombia
Catálogo de germoplasma de frijol/editor, Rigoberto Hidalgo. - Cali, Colombia: Centro Internacional de Agricultura Tropical, Dic. 1984. - p.: ilus.
ISSN: 0120-6265
,. Frijol- Germoplasma - Catálogos y colecciones. l. Hidalgo, Rigoberto. 1/. Centro Internacional de Agricultura Tropical.
La Unidad de Recursos Genéticos es responsable
del manejo del germoplasma de los principales progra
mas del CIAT: frijol, yuca y pastos tropicales.
Esta responsabilidad incluye la. colección, evalua
ción, conservación e intercambio, así como la docu
mentación y distribución de la información acerca
del germoplasma de dichos cultivos. Las actividades
de la Unidad aseguran que los valiosos, y a veces
escasos recursos genéticos, sean protegidos cuidado
samente y a la vez, plenamente utilizados tanto
en el CIAT como en el resto del mundo.
The Genetic Resources Unit is responsible
for the management of
ties of CIAT: beans,
germplasm of the main commodi
cassava and tropical pastures.
This responsibility includes the collection, evalua
tion, conservation and exchange, as well as the
documentation and distríbution of informatíon related
to those crops. The activítíes of the Unit ensure
that valuable and sometimes rare genetíc resources
will be carefully protected, but at the same time,
fully utilízed both in CIAT and throughout the world.
jii
iv
PERSO~AL DE LA @~I~AD ~E RECURSOS GE~ETICOS (PEMSOMMEL)
CIENTIFICO PRINCIPAL (SENIOR STAFF)
William M. Roca, Ph.D. (Acting Head)
CIENTIFICO POSDOCTORADO (POST-DOC FELLOW)
László M. Szabados, Ph.D.
ASOCIADO DE INVESTIGACION (RESEARCH ASSOCIATE)
Rigoberto Hidalgo, M.Sc. - Frijol (Beans)
ASISTENTE DE INVESTIGACION (RESEARCH ASSISTANT)
Hember Rubiano, Ing. Agr. - Frijol (Beans) Javier Belalcázar, Ing. Agr. - Pastos (Pastures) Javier Narváez, Ing. Agr. - Cultivo Tejidos (Tissue Culture) Javier Beltrán, Biol. - Cultivo Tejidos (Tissue Culture) Graciela MafIa, Biol. - Cultivo Tejidos (Tissue Culture) Isabel Natalia Salas, Biol. - Sanidad Semillas (Seed Health)
EXPERTO 1 (EXPERT 1)
Orlando Toro, Tec. Agr. - Frijol (Beans) Julio Roa, Biol. - Cultivo Tejidos (Tissue Culture)
TECNICO 1 (TECHNICIAN 1)
Julieta Ortiz - Procesamiento Datos (Data Processing) Raul Reyes, Biol. - Cultivo Tejidos (Tissue Culture)
SECRETARIA (SECRETARY)
Helga Dierolf
PERSONAL ASOCIADO (ASSOCIATE PERSONNEL)
Miguel Halle, Ph.D. - CIRF (IBPGR)- Coordinador Regional América Latina (Regional Coordinator Latín America)
La Unidad de Recursos Genéticos es responsable
del manejo del germoplasma de los principales progra-
mas del CIAT: frijol, yuca y pastos tropicales.
Esta responsabilidad incluye la. colección, evalua
ción, conservación e intercambio, así como la docu
mentación y distribución de la información acerca
del germoplasma de dichos cultivos. Las actividades
de la Unidad aseguran que los valiosos, y a veces
escasos recursos genéticos, sean protegidos cuidado-
samente y a la vez, plenamente utilizados tanto
en el CIAT como en el resto del mundo.
The Genetic Resources Unit is responsible
for the management of germplasm of the main commodi
ties of CIAT: beans, cassava and tropical pastures.
This responsibility includes the collection, evalua-
tion, conservation and exchange, as well as the
documentation and distribution of information related
to those crops. The activities of the Unit ensure
that valuable and sometimes rare genetic resources
will be carefully protected, but at the same time,
fully utilized both in CIAT and throughout the world.
iii
INTRODUCCION vii
INTRODUCTION ix
Status de la colecci6n de Phaseolus xi
Procedencia del germoplasma de f. vulgaris xii
Distribuci6n del germoplasma de P. vulgaris de acuerdo xiv a color y al hábito
Descriptores xv
Status of the Phaseolus collection xxi
Source of the germplasm of f. vulgaris xxii
Distribution of the germplasm of f. vulgaris xxiv according to color and growth habit
Descriptors xxv
Lista secuencial por No. G 1
Sequential listing by G number 1
Grupos de color y tamaño de semilla 102
Groups by color and seed size 102
Blancos (White) 102 Grandes (Large) 102 Medianos (Medium) 112 Pequeños (Small) 126
Crema-beige (Cream-beige) Grandes (Large) Medianos (Medium) Pequeños (Small)
Amarillo (Yellow) Grandes (Large) Medianos (Medium) Pequeños (Small)
Café-marr6n (Brown-marron) Grandes (Large) Medianos (Medium) Pequeños (Small)
Rosado (Pink) Grandes (Large) Medianos (Medium) Pequeños (Small)
-~ "
.-~-- . ...; '~---_ ....
143 143 158 174
183 183 189 202
207 207 209 213
218 218 221 224
v
vi
Rojo (Red) Grandes (Large) Medianos (Medium) Pequeños (Small)
Morado (Purple) Grandes (Large) Medianos (Medium) Pequeños (Small)
Negro (Black) Grande (Large) Medianos (Medium) Pequeños (Small)
Otros (Other) Grandes (Large) Medianos (Medium) Pequeños (Small)
229 229 234 241
254 254 259 264
267 267 269 280
316 316 316 317
UlTIlODlIJCCIOllll
Dentro del sistema de Centros Internacionales, CIAT tiene la responsabilidad mundial para la colección y preservación del germoplasma de frijoles del género Phaseolus. Siete años después de haber asumido esta responsabilidad, la colección de germoplasma del CIAT tiene más de 33000 accesiones (ver tabla), la cual abarca las cuatro especies cultivadas, sus ancestros silvestres, así como las especies silvestres no cul ti vadas. Este germoplasma se ha obtenido a través de donaciones de los bancos de germoplasma de Programas Nacionales y de expediciones a los centros de origen y de dispersión. También se incluyen los materiales más prornisorios de las progenies resultantes de los esfuerzos de hibridación del Programa de Mejoramiento de Frijol.
El rango de adaptación de f. vulgaris como cultivo va desde O hasta 3000 m.s. n.m. y desde O a 42 0 latitud N y S, lo cual ha generado una gran variabilidad genética. Hallazgos arqueológicos en Mesoamérica así como la disponibilidad de un gran número de formas silvestres ancestrales y de formas silvestres no cultivadas del género en esta región, sugieren al área de México/Guatemala como el centro de origen del frijol común. Sin embargo, desde las expediciones de Colón, el frijol común se ha dispersado geográficamente y eventualmente ha sido adaptado a muchas nuevas áreas fuera de América, dando como resultado la creación de centros secundarios de dispersión, los cuales en muchos casos tienen cerca de 400 años. El CIAT continua asignando alta prioridad a los esfuerzos de colección en estas áreas, en estrecha colaboración con el Consejo Internacional de Recursos Fitogenéticos (CIRF), dando especial énfasis en aquellas áreas con avanzada erosión genética y con alta probabilidad de encontrar razas nativas y formas silvestres de la especie.
Este catálogo incluye más de 16000 accesiones de formas cultivadas de f. vulgaris que han sido incrementadas por la Unidad de Recursos Genéticos del CIAT. Debido al tamaño de esta colección sólo se ha incluído información pasaporte clave, así como características de semilla y del hábito de crecimiento. Hay tres secciones principales: a) Breve descripción de la colección y definición de descriptores; b) Listado secuencial de los número G, que dan la clave de tipo de semilla; c) Listado de las accesiones agrupadas por color y tamaño de semilla.
Semillas (pequeñas cantidades) y/o información adicional sobre la caracterización del germoplasma incluido en este catálogo, están disponibles bajo pedido, escribiendo a la siguiente dirección:
R. Hidalgo Unidad de Recursos Genéticos CIAT Apartado Aéreo 67-13 Cali, Colombia Suramérica
vii
UTJlODUCYIOW
Wíthin the International Center' s system, CIAT has global responsability for collectíon and preservatíon of germplasm oi beans of the genus Phaseolus. Seven years after formally assumíng the responsabílíty, the CIAT germplasm collection contaíns more than 33000 accessions (see table), embracíng the tour cultívated species, theír wild ancestors and the wíld non-cultivated species. This germplasm has been obtained through donations from national germplasm banks and expeditions to the centers oi origin and dispersion. Also included in the bank are the most promising progenies resulting from the hybridization efforts at CIAT.
Adaptation of f. vulgaris as a crop ranges from O to 3000 m.a.s.l., and from O to 42° latitude N and S, generatíng a broad genetíc varíability. Archeological findíngs in Mesoameríca and the availability of a large number of wild ancestral and wild noncultivated forms of the genus there, suggest that Mexico/Guatemala area is the center of origin f or the common bean. However, since the expeditíons of Columbus, the common bean was geographícally díspersed and thus has been adapted to many new areas outsíde the Amerícas resulting in several secondary centers of dispersion, which, ín many cases are about 400 years old. Active collection efforts for f. vulgaris in those areas contínue to be assígned high príority by CIAT in clase collaboratíon with the International Board for Plant Genetic Resources (IBPGR) with special emphasís being placed on areas wíth advanced genetíc erosion and wíth a hígh probabilíty of finding land races and wild forms of the cultivar.
This catalog includes more than 16000 accessions of cultivated forms of f. vulgaris, which have been increased by the Genetic Resources Unit of CIAT. Due to the síze of this collection only key passport information as well as seed type descriptíon and growth habit are included. There are three main sections: a) brief description of the collectíon and definitíon of descriptors¡ b) Sequential G numbering which addresses to the seed type¡ c) list of accessíons grouped by seed color and seed size.
Seeds (sma11 quantities) or additional ínformatíon on characterization of the germplasm listed in thís catalog can be avaílable upon request by wríting to:
R. Hídalgo Genetic Resources Unit CIAr Apartado Aéreo 67-13 Cali, Colombia South America
ix
STATUS DE LA COLECCION DE FRIJOLES PHASEOJ.US ALMACENADOS EN LA
UNIDAD DE RECURSOS GENETICOS DEL CIAT (Junio, 1984)
E s p e c i e s
Cultivadas y silvestres ancestrales
P. vulgaris
P. lunatus
P. coccineus
f.. acutifolius
Silvestres no-cultivadas
*
P. anisotrichus, f.. filiformis,
P. galactoides, f.. microcarpus,
P. metcalfei, f.. pedicellatus,
P. polystachius, f.. parvulus,
P. ritensis, P. wrightii
TOTAL
Semilla disponible.
No. de accesiones
Introducidos
29552
2410
1083
166
84
33295
Incrementados*
16799
700
220
166
39
17924
xi
xii
PROCEDENCIA DEL GERMOPLASMA DE P. VULGARIS* INCLUIDO EN ESTE
CATALOGO
REGION-PAIS
Norteamérica
Estados Unidos
Subtotal
Centroamérica
Costa Rica El Salvador Guatemala Honduras México Nicaragua
Caribe
Belice Cuba
Subtotal
República Dominicana Haití Jamaica Puerto Rico Islas Guadalupe
Subtotal
Sur américa Zona Andina
Bolivia Colombia Chile Ecuador Perú Venezuela
Subtotal
Suramérica Zona No-Andina
Argentina Brasil Uruguay Guyana
Subtotal
ACCESIONES
5084
5084
469 98
1315 1147 1118
6
4153
6 4 7
26 3
37 1
84
86 934 127 355 844 311
2657
77 531
10 2
620
* No se incluyen formas silvestres.
PORCENTAJE
30.2
24.7
0.5
15.8
3.7
REGION-PAIS ACCESIONES PORCENTAJE
Europa
Bulgaria 10 Belgica 32 Francia 576 Alemania (Este) 199 Grecia 15 Hungria 126 Italia 621 Holanda 885 Portugal 279 España 71 Unión Sovietica 38 Inglaterra 755
Subtotal 3607 21.4
Africa
Kenia 15 Malawi 125 Nigeria 12 Suráfrica 8 Zambia 12
---Subtotal 172 1.0
Asia - Oceanía
Australia 22 China 2 Japón 344 Nueva Zelandia 24
Subtotal 392 2.3
TOTAL 16769 100.0
xiii
l. No. CIAT = Número CIAT
Número CIAT o número de germoplasma es el número asignado
a la accesión original en el banco de germoplasma de la Unidad
de Recursos Genéticos del CIAT, cuando la semilla ha sido
incrementada~ Este número consta de la letra G seguida de
cinco dígitos. En algunos casos hay una letra después de
los dígitos CA, B, C, etc.). Esta letra corresponde a separa-
ciones justificadas de la accesión original.
2. IDENTIFICACION = Identificación
Es el nombre local o varietal de la accesión o el número
de registro del donante de la semilla. Este nombre se registra
tal como se recibe de la fuente donadora y sólo se hacen
correcciones en nombres de materiales muy conocidos.
3. REGLOCAL = Registro Local
4.
Corresponde al registro de la accesión usado por el
país, el investigador o el colector donante. Este registro
puede ser numérico o alfabético-numérico. En algunos casos
puede reemplazar a la identificación, cuando esta no existe
o se desconoce.
ORI Origen
En general, corresponde al país que previamente envió
la . ,
acceSl.on al pals donante de la semilla al ClA T • Sin
embargo, si se conoce con certeza el país donde se colectó
o mejoró el material, este se registra sin tener en cuenta
el resto de la información. En algunos materiales el "origen"
no aparece debido a que la información disponible no es sufi-
ciente para determinarlo. Los códigos para los países son
los mismos que se usan para procedencia.
xv
xvi
5. PROCED = Procedencia
Es el país donante de la semilla a CIAT. Los siguientes
códigos son los usados para los países.
AFG = Afganistán
AGA = Argelia
AGL Angola
ARA = Arabia
ARG = Argentina
ATL = Australia
BGA = Bulgaria
BLG = Bélgica
BLV = Bolivia
BND = Burundi
BZE = Belice
BZL = Brasil
CBA = Cuba
CDA = Canadá
CHI = China
CHN = Taiwan
CLB = Colombia
CLE = Chile
CME = Camerún Británico
CMF = Camerún Francés
CMR = Camerún
CNG = Congo
CPR = República Popular China
CPS Chipre
CRA = Costa Rica
CSR = Checoslovaquia
DMK = Dinamarca
DOM = República Dominicana
ECD = Ecuador
EGT = Egipto
ELS = El Salvador
ETP = Etiopía
FRC = Francia
GAM = Gambia
GDR = Alemania (Este)
GFR Alemania (Oeste)
GMN = Alemania
GRC Grecia
GTA = Guatemala
GUY = Guyana
HDR = Honduras
HGY = Hungria
HTI = Haití
IDS = Indonesia
IGU = Islas Guadalupe
IND = India
IRN = Irán
ITL = Italia
JDN = Jordania
JMC = Jamaica
JPN = Japón
KOR = Corea
KYA = Kenia
LBN = Líbano
MEX = México
MWI = Malawi
NCA = Nicaragua
NGA = Nigeria
NLD = Holanda
NPL = Nepal
NWY = Noruega
NZL = Nueva Zelandia
PER = Perú
PKS = Pakistán
6.
PLD = Polonia SWL = Suiza
PRG = Paraguay TKY = Turquía
PRI = Puerto Rico TZA = Tanzania
PTC = Portugal UGA Uganda
RHO = Rodesia URG = Uruguay
RMN Rumania USA = Estados Unidos
RWD = Rwanda UTK = Inglaterra
SAF = Suráfrica VNZ = Venezuela
SAR = República Arabe Siria YEM = Yemen
SDN = Sudán YUG Yugoslavia
SPN = España ZBA = Zambia
SSU Unión Sovietica ZRE Zaire
SWD = Suecia
CP = Color Primario de Semilla
CS = Color Secundario de Semilla
Se consideran 9 grupos que corresponden a 7 familias
de colores. Dentro de cada familia se incluyen todos los
rangos de tonalidades, tratando hasta donde sea posible de
disminuir la clasificación subjetiva de los colores. Sólo
se tienen en cuenta el color primario, que es el predominante,
y el color secundario de la semilla cuando ésta presenta dos
o más colores. Tanto el color primario como el secundario
se toma de semilla recientemente cosechada y seca.
Grupos de Familias
Bl = Blanco
Cr Crema-beige
Tipo varietal Panamito Blanquillo Arroz Navy Bean Small White Great Northern Alubia Coscorrón Caballero
Carioca Mulatinho Bico de Ouro
País Perú, Ecuador Colombia Chile Estados Unidos Estados Unidos Estados Unidos Argentina Chile Perú
Brasil Brasil Brasil
xvii
niii
Am = Amarillo
Cf = Café-marrón
Rs = Rosado
Rj = Rojo
Mr = Morado
Ng = Negro
Ot = Otros
Cargamanto Pinto Ojo de Cabra Bayo
Canario Azufrado Enxofrao Liborino Jalo Amarelo
Ricopardo Redlands Chumbinho
Rosinha Rosita Flor de Mayo Miss Kely Red Kidney
Andino
Calima Pompadour Zamorano
Sangre toro Roxinho
Porrillo Jamapa Turrialba lCA-Pijao Rio Tibagi Caraota
Tórtolas Burros
7. TAMANO = Tamaño de Semilla
Colombia México México México, Perú, Chile,
Ecuador
Perú, México México Brasil Colombia Brasil Brasil
Brasil Australia Brasil
Brasil México México Jamaica, Belice EEUU, Jamaica, Belice
Panamá Colombia
Colombia República Dominicana Honduras
Colombia Brasil
El Salvador México Costa Rica Colombia Brasil Venezuela
Chile Chile
Es el peso en gramos de una muestra randomizada de 100
semillas. Sin embargo, como forma práctica de clasificación
se han formado 3 grupos principales de la siguiente manera:
Pequeños: hasta 25 g/IDO semillas
Medianos: entre 25 y 40 gllOO semillas
Grandes: desde 40 g/IDO semillas
8. FS = Forma de Semilla
Como en el caso de los colores, el criterio es formar
grupos con similitudes en cuanto a la forma se refiere. La
forma se determina observando la semilla de lado.
00 O
00
1 . redondeada 2. elongada
9. BLLO = Brillo de la Semilla
Se toma en semillas secas
1 = Opaco
2 = Intermedio
3 = Brillante
10. MEZR = Mezcla
3. arriñonada
Se refiere a la presencia de semillas con color, brillo,
forma y tamaño diferentes a la semilla que está en mayor pro-
porción dentro de la accesión original. Esto es frecuente
en materiales que han sido colectados en expediciones o en
poblaciones provenientes de programas de mejoramiento. Se
tienen 3 criteríos para clasificar una mezcla.
MM = mezcla mecánica
MS = mezcla segregante
2M mezcla ,
= mecanica y segregante
xix
xx
11. HABT = Hábito de Crecimiento
Es un concepto morfoagronómico definido como el resultado
de la interacción de varios caracteres de la planta que deter-
minan su arquitectura final. Estos caracteres son: Tipo
de desarrollo de la parte terminal del tallo (determinado
o indeterminado), número de nudos en el tallo, longi tud de
entrenudos, aptitud para trepar y patrón de ramificación.
Se han definido 4 prototipos principales de hábito así:
1 = Arbustivo determinooo
2 = Arbustivo indeterminado
3 = Indeterminado postrado con una o varias guías largas,
pero con ninguna o muy poca aptitud para trepar
4 = Indeterminado con guía larga y trepadora
STATUS OF THE PHASEOLUS BEAN COLLECTION HELD AT THE
GENETIC RESOURCES UNIT OF CIAT (June - 1984)
No. of accessions
Species
Cultivated and wild ancestral
f· vulgaris
f· lunatus
P. coccineus
f· acutifolius
Wild non-cultivated
P. anisotrichus, f. filiformis,
P. galactoides, f. microcarpus,
f. metcalfei, f. pedicellatus,
P. polystachius, f. parvulus,
f. ritensis, f. wrightii
TOTAL
* Seed available.
Introduced
29552
2410
1083
166
84
33295
Increased*
16799
700
220
166
39
17924
xxi
xxii
SOURCE OF THE GERMPLASM OF P. VULGARIS* INCLUDED IN THIS CATALOG
REGION-COUNTRY
North America
United States
Sub total
Central America
Costa Rica El Salvador Guatemala Honduras Mexico Nicaragua
Caribbean
Belize Cuba
Subtotal
Dominican Republic Haiti Jamaica Puerto Rico Guadalupe Islands
Sub total
South America Andean Zone
Bolivia Colombia Chile Ecuador Peru Venezuela
Subtotal
South America Non-Andean Zone
Argentina Brazil Uruguay Guyana
Subtotal
* Wild forms are not included.
ACCESSIONS
5084
5084
469 98
1315 1147 1118
6
4153
6 4 7
26 3
37 1
84
86 934 127 355 844 311
2657
77 531
10 2
620
PERCENTAGE
30.2
24.7
0.5
15.8
3.7
REGION-COUNTRY ACCESSIONS PERCENTAGE
Euro12e
Bulgaria 10 Belgium 32 France 576 Germany (East) 199 Greece 15 Hungary 126 Italy 621 Holland 885 Portugal 279 Spain 71 Soviet Union 38 United Kingdom 755
Subtotal 3607 21.4
Africa
Kenya 15 Malawi 125 Nigeria 12 South Africa 8 Zambia 12
Subtotal 172 1.0
Asia - Oceania
Australia 22 China 2 Japan 344 New Zealand 24
Subtotal 392 2.3
TOTAL 16769 100.0
xxiii
" " <. DISTRIBUTION OF THE GERMPLASM OF P. VULGARIS* ACCORDING TO COLOR AND GROWTH HABIT
CREAM BROWN WHITE BEIGE YELLOW MAROON PINK RED PURPLE BLACK OTHERS TOTAL
HABIT 1
Frequency 1107 594 802 190 172 322 283 270 10 3750
Percentage 34.7 18.8 42.1 23.4 21.9 16.4 28.2 7.0 9.2 22.4
HABIT II
Frequency 144 279 80 97 66 128 95 1336 7 2232
Percentage 4.5 8.8 4.2 12.0 8.4 6.5 9.5 34.7 6.4 13.3
HABIT III
Frequency 976 1182 462 218 295 1056 298 1121 34 5642
Percentage 30.6 37.5 24.2 26.9 37.6 53.9 29.7 29.1 31.2 33.6
HABIT IV
Frequency 815 960 502 262 226 396 281 1055 50 4547
Percentage 25.6 30.4 26.3 32.3 28.8 20.2 28.0 27.4 45.9 27.1
WITHOUT HABIT
Frequency 144 138 60 44 26 58 45 70 8 593
Percentage 4.5 4.4 3.1 5.4 3.3 3.0 4.5 1.8 7.3 3.5
TOTAL
Frequency 3186 3153 1906 811 785 1960 1002 3852 109 16764
Percentage 19.0 18.8 11.4 4.8 4.7 11.7 6.0 23.0 0.7 100.0
* Wild forms are not included
l. No. CIAT = CIAT's Number
CIAT's number or germplasm number is the number assigned
to the original accession in the Genetic Resources Unit when
the seed has been increased. This number has five digits
preceded by the letter G. In sorne cases a letter (A, B, C,
etc.) follows the five digits; this letter corresponds to
justified separations from the original accession.
2. IDENTIFICATION = Identification
It is the local or varietal name of the accession, or
the local register used by the seed donor. It is recorded
as originally received, and corrections are made only on names
of very well known materials.
3. REGLOCAL = Local Register
It corresponds to the register number used by the natio
nal institution, scientists or collector donating the seeds.
This local register can be either numerical or alpha-numerical.
In some cases it could be used as ider.tification when the
latter does not exist or is unknown.
4. ORI = Origin
In most instances it is the country from which the acces
sion was obtained before it was donated to CIAT. However,
if it is known where the material was bred or collected, su eh
country is recorded as "origin" ignoring the different previous
sources. When the information available is not enough to
determine the "origin'! it is not included. The codes for
countries are the same ones used for the source.
5. PROCED = Source
It is the seed donor's country. The following codes
xxv
are used for countries.
AFG = Afganistan GFR = Germany (West)
AGA = Algeria GMN = Germany
AGL = Angola GRC = Greece
ARA = Arabia GTA Guatemala
ARG Argentina GUY Guyana
ATL = Australia HDR Honduras BGA = Bulgaria HGY = Hungary
BLG = Belgium HTI Haiti
BLV = Bolivia IDS = Indonesia
BND = Burundi IGU = Guadalupe Islands
BZE Belize IND = India
BZL = Brazil IRN Iran
CBA = Cuba ITL Italy
CDA = Canada JDN Jordan
CHI = China JMC = Jamaica
CHN = Taiwan JPN Japan
CLB = Colombia KOR = Korea
CLE Chile KYA = Kenya
CMB = Cameroons, British LBN = Lebanon
CMF = Cameroons, French MEX = Mexico
CMR = Cameroon MWI = Malawy
CNG Congo NCA = Nicaragua
CPR = People's Republic of China NGA Nigeria
CPS = Cyprus NLD = Holland
CRA = Costa Rica NPL Nepal
CSR = Czechoslovakia NWY Norway
DMK = Denmark NZL New Zealand
DOM = Dominican Republic PER Peru
ECD = Ecuador PKS = Pakistan
EGT = Egypt PLD = Poland
ELS = El Salvador PRG = Paraguay
ETP = Ethiopia PRI Puerto Rico
FRC = France PTC = Portugal
GAM = Gambia RHO = Rhodesia
GDR = Gennany (East) RMN = Rumania
xxvi
RWD Rwanda
SAF ~ South Africa
SAR Syrian Arabic Republic
SDN ~ Sudan
SPN Spain
SSU ~ Soviet Union
SWD Sweeden
SWL Switzerland
TKY ~ Turkey
TZA ~ Tanzania
6. CP ~ Primary Seed Color
CS ~ Secondary Seed Color
UCA ~ Uganda
URC ~ Uruguay
USA ~ Uníted Sta tes
UTK ~ United Kingdom
VNZ ~ Venezuela
YEM ~ Yemen
YUC ~ Yugoslavia
ZBA Zambia
ZRE ~ Zaire
There are nine color groups deri ved from seven color
families. In order to avoid a subjecti ve classification of
color, an attempt has been made to include the widest spectrum
possible of shades wi thin each color. The primary color is
the predominant seed coat color; if the seed has more than
one color, only the secondary is recorded.
from dry seeds recently harvested.
Color is taken
Fami1y Graups
BI ~ White
Cr ~ Cream-beige
Varietal seed types
Panamito Blanquillo Arroz Navy Bean Small White Creat Northern Alubia Coscorron Caballero
Carioca Mulatinho Bico de Ouro Cargamanto Pinto Ojo de Cabra Bayo
Cauntry
Perú, Ecuador Colombia Chile United States United States Uníted States Argentina Chile Perú
Brazil Brazil Brazil Colombia Mexico Mexico Mexico, Peru, Chile,
Ecuador
xxyii
xxviii
Am = Yellow Canario Azufrado Enxofrao Liborino Jalo Amarelo
Cf = Brown-maroon Ricopardo Redlands Chumbinho
Rs = Pink Rosinha
Rj = Red
Mr = Purple
Ng = Black
Ot = Others
7. TAMANO = Seed Size
Rosita Flor de Mayo Miss Kelly Red Kidney
Andino
Calima Pompadour Zamorano
Sangretoro Roxinho
Porrillo Jamapa Turrialba lCA-Pijao Río Tíbagi Caraota
Tortolas Burros
Peru, Mexíco Mexico Brazil Colombia Brazil Brazil
Brazil Australia Brazil
Brazil Mexico Mexico Jamaica, Belize USA, Jamaica, Belize,
Panama Colombia
Colombia Dominican Republic Honduras
Colombia Brazil
El Salvador Mexico Costa Rica Colombia Brazil Venezuela
Chile Chile
It is the weight of a sample of 100 seeds taken at ran
domo However. as a practical mean of classification three
main seed sizes have been established, as follows:
Small:
Medium:
Large:
les s than 25 g/lOO seeds
between 25 g and 40 g/lOO seeds
larger than 40 g/lOO seeds
8. FS = Seed Shape
As with the color grouping, the criteria for seed shape
is also to conform groups with close in shape similarity:
00 O
l. rounded
9. BLLO = Brilliance
00
2. elongated 3. kidney type
It is taken on dry seed recently harvested.
1 = Opaque
2 = Intermediate
3 = Brilliant
10. MEZR = Mixture
It refers to the presence of seeds with color, brilliance,
shape or size different from the seed type of highest percen
tage in the original accesion. This is frequent in materials
collected in expeditions or in populations produced in breeding
programs.
mixture:
Three criteria have been defined to classify a
MM = mechanical mixture
MS = segregating mixture
2M = mechanical and segregating mixture
11. RABT = Growth Rabit
It is a morphoagronomic concept defined as the result
of the interaction of several characters that determine the
final plant architecture. These characters are: type of growth
of the stem (determinate or indeterminate), number of nodes
on the stem, length of the internodes, climbing abili ty and
branching pattern. Four prototypes of growth habit have be en
defined as follows:
xxix
xn
1 = Bush, determinate
2 Bush, indeterminate
3 = Indeterminate prostrate with lateral branching well
developed, but low or nil climbing ability
4 = Indeterminate with long guide and high clímbing
ability
...
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2 2
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IVT-141J7 NLO NLO lYT-73127 F~C NLD lvf-7220Q USA NLD IVT-75270 GF~ ~LD lVT-75191 NLD NLO
~
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2 G0761b AJCHERS LEBENER ~ErSTERwEk COR
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1 IG076~7 MARVANYSlE~U G07669 MIGNON G07690 MOOERNA
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G0771. SCHL_C~TSCHWERT EXTRA LANG 1 IG07725 TERRAS VE~aeSSEATE PRINlEe 1 G07726 TQP NQTCH GOLDE N ~Ax
1 G07739 Z_AAN'S RESISTENTI4 1 G07921 DE" 38-3 GO 7924 OE. 4. 1 GC7926 DE 43 3 GQ7929 ALUBIA (POBLACION COMVNJ
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2 2
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eTA t.:.TA (.TA C,TA GTA. t.,..TA C.TA ~TA
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GT¡k {,TA
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GTA (;JA GTA (,T~
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(;ToI\ \...TA G1"A -erA (;1'"" GTA GTA (,.TA
GTA GTA
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I 1 1 I 1 I
2 3 2 3 2 2 2 3 2 3
2 J 3
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G108,<¡7 CUATE~.LA. 137'1 C.108a~ GVATEMALA 13~O
GIOSR9 GUATEMALA 1382 GI0890 GVATEN"'L' 13iB G10895 GU~TE~ÁLA 1399
4 ~10901
.. G,1090¿
4 GI0916 4 G10925 4 G10933
GU~TtflllALA 140.3
Gl,J.<t.TEMALA 140'
GUATFMALA 14)\ Gu .... rEf'lALA 1499 GUAT¡::: .... At.A 1513
". G10939 GUATFMALA 1520 4 Gl10Ab Cu~qlTO
4 Gl1126 CHATIFA DE ETAMP 4 Gll13~ JANTAR 4 GI111? KúNS(RVNAJA
4 Glll?Q P~PITA
4 Glll:q S AHIC.. A
4 1'..1116Q SI-QOKQSTUCKDvAJa..
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G 11224
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Gl1271 GU.278 <111279 Gl121'\O
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lvt 721.
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GTIt-1319 <;T A- 1380
GTA.-1362
GT"-1383 GT 1\-1369
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GTA-1431
C.TÁ-1499
GT¡l-1513
GoTA GTA
GrA GTÁ GTA GTA GTA GTA (iTA GTA
G,TA GTAo
t;TA GTA <iTA GTA
GTAo GT .... GTA eTA
6TA-1520 GT. GTA DG078/159 NEX MEX !Vl-773Q7 SSU NlO IVT-71423 S5U NLO JVT-17426 ssu NLD
IVT-77~65 NLD NLO lvT-7710b ssu NLO !yT-71106 SSU NLO lvr-77106 ssu NLO lVT-11b94 NLO NLD
JVT-71~Q5 PLO NLD [VT-176Q6 GFR ~LD Ivr-71914 USA NLD lVT-779~4 ~FR NLO IVT-71948 CSR NLO
IVT-77'~Só NLO NL() IVT-77969 NLO NLO IVT-779Q4 NLO NLO !VT-77100 GFR NLO IVT-17l02 GFR NLO
lVT-17102 GFR NLD ¡~T-77104 GFP NLD tVT-77¡05 NLO NLU lVT-73J21 NLO NLO [Vr-76064 ~LD NLO
Ivr-71217 ~LO NLO IVT-77302 USA NLO IVT-77303 USA NLO Ivr-77645 GFR NLD R$ 1vT-77844 GF~ NLO
2 2 2 3 2 3 2 3 2 3
2 3 2 3 2 3 2 3
2 3
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,-----:------------------I NO. ICJAr I
IOENTIFICACION 'REGLOCAL
I I
O I p I Q I R le
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Glt339 AGVASCALIENTES 19-1 Gtt455 VEPACAUz 6-2 Gllb02 ~lJ644 BOCA A~ARILLA
Gl1854 WALCHERS~ wlTTE CLPTI
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G12.~8 8LANCO TIPO ÑEP (BOLICHE) G1249S ~CLA 63 ! G12613 PA~A~lTO GRANDE G12725 L.~3007-(MC.~M.-HC3)
G12737A ~720IC-17-. FARLY SELECT.
G1]1~a P[LA~58E~G
G13159 H::f'SUS G13182 T~~ASH[~OTEBG
G1JIR3 UN(HowN tus., 61-3223 UN<..NQ'IIN-2
(.13274
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G\3285
Gl.3286
GJ.3287 G1326A Gl J269 G13290 G13291
C.13¿Q9
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GI3.JOl G 1330 l (, l 3304
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NLO NLD
"'LO NLO hLD tilLO USA USA USA USA
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CAT-0923- ECO CAT-ln74 ECO COMP.OIO PER P!;R
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COO-1I02 USA JPN COO-1250 JPN JPN
coo- L403
COO-21Q5
COD-'?lO 'J
COO-2111 COO-2112
COD-2111 COO-2114 COO-211 'S COO-2.116 COO-2117
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UNKNOwN ¿58l UNt(HOW,,", 2591 VNKNOWN 2592 UNI<.NOWN 2593 UNKNOwN 25q4
[f<Ató RE'ZAYEH ql LEBANQN IRAN MIN./t31 312 GREECE IRAN ~lN/l45 3~6 UNKNQViN 5-14 VN+(NO .. N SIC)
COO-3IQ7 jPN JPN COD-311. ros JPN COO-3116 lNO JPN caO-6109 jPN JPN COP-6110 JPN JPN
CGD-eI17 VTK JPN PHA0880~7 úDR ~-16452 MEX MEX RS VP-035 ~EX #!IEX lH60-0176 Z8A [TL ~~
, 2
1 2 J 2
1 3 2 2 1 2 2 2 2 2
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VAV3131-a ARG ~RG RJ 2 2 FRC-0093 R~N FRC 1 2 FRC-0636 NLO FRC 2 1 fRC-08ól FRC RS 2 1 FRC-0895 U5~ FRC 2 2
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PI416600 Pl4t66Q.]
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pr"16609 Pt4l~610
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P1416648 PJ4l665Z P1416657 P1416675 P1416677
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USA 2 us~ Z USA 2
U-SA 2
USA 2 USA 2 USA 1 2
USA 1 2
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G14146 UNKNQWN 532 G141_a CVPRUS'Y~R MORPHIAICA 592 G14150 LEBA~ON 646 G141S9 rURK~Y 1196 G14168 TURK~Y'5el'2 2440
G142.82 G144J o G14415
G14471 G14S01
307
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GI4SJ6 8.VE~DURÁ DE SH JERONJ_O G14S44 "OINOA &14545 AFRA G14S47 ASTAI=:! G14549 eovT"DE
(;14551 EAGLE G14553 GA!..A G14S58 KINGCAOSS .ACHS G14S59 LAOueO G14S61 MINISEL
G14562 Ol.wADE GJ4S6.1 ROLANDO G14564 SAS¡;'¡ESCO G14565 SLIM6EL G14S66 SA)(ANOVA
G14S67 51· ...... ro G14S68 T~~OERETTE
Gol 4569 VALGPEEN Gl*575 GQ G14577 GCLD~RONt
GIlt581 RHF::INLANO G14S83 WE18E FELD60HNE Gl4S68 DELIKATA GI4598 GUAT=MALA 252 G14tiOl GUATEMALA 616
Gll!.602 GI"607 G14b12 GI4613 G141')14
GUAf::::II4Al.A 62')
GUA TEIofALA 768
Ohf"GQN se A:
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P1416678 USA
PI416680 CPS USA PI41668. LBN USA
PI416702 TKY USA Pl416720 T~Y USA
1 2 3 2 1 2 1 2 2 2
MWI CF 1 2 e.t..osaSA C.L.0636 PHA1056'7 pi 289338
SPN Uf K .3
SPN UTK 1 3
GDA 2 3 HGY USA 2 .]
HDR-2502 HDR HO~ IVT-74118 NLO NLD IVT-73195 GUR NLO IVT-73108 FRC NLD ¡VT-73166 NLO NLO
IVT-73201 USA NLO IVT-73119 FRC N~O J Vr-71'167 NLO Nl.O IVT-7~~50 NLO NLO lVT-77155 NLO N~O
lVT-72261 NLD NLD JV7-7518~ NLD NLO lVT-74185 NLD NLD IVT-77212 NLD NLo lVT-73164 GOR NLD
JVT-74198 NLD NLo IVT-73266 USA NLO Ivr-73299 USA NLD PHA0691,7 GDR PHA0344,1 GD~
PHA02b3,7 GOR PHA0573,7 GO~
PH41095/7 GOR
(j,TA-252 GTA GTA GTA-616 GTA eT'"
GTA.-625 GTA ("TA GTA-168 GT ... &TA
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2 3 3
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1 2
1 2 2 3 2 2 2 2
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4 3 4 1 3
G14ó27 G14b30 Gllj.ó31 Gl4632 G14633
MOLIMA ORELLA PA:ERESCO REFLEX ROVNOUP
3IG1"6.10 TRfASURE l Gt4631 TROVATO t G14639 WALCHERSE WITTE
G146.7 66-779 G14674 ALU8lA GRANOE
(014677 ESPAÑA 5 1 IG14680 BLANQUILLO 1 G'.ó89 WHITE cot:o 1 G14692 SAPoSTELA OWARF
G14700 SELECC10N DE MEZCLA Xl
G14703 CACHANUN01NHO VII 1 IG14711 VAQUITA lO,ROQUE,GTO. l G14727 ST.A 1 G14734 ALLE FRUHESTE wUSSE t G147l5 AMATHYST
1 'Gl4738 A.R.22 G14148 BOLITAS
1 IG14750 SBL 274 • Gl41s2 CEALI 4 Gl.157 COSLA ~ (OBLAN
4 ~14771 coco TQUkONNAIS NA1N 1 Gl4774 COLINA J GJ477ó COMBRIO 4 G14779 EXPO RUSSE 4 G14781 GALLATIN 50
• 4 1 1
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G. N. U. l. 1130 G. N. OIA""f:.RJQUE LE GR 1GNOR~IS HYSCORE Il'\f.PUISA.BLE
IVT-77514 NLO NLO IVT-77576 FRC NLD JVT-77S78 USA NLO lVT-77S85 USA NLO IVf-77667 NLD NLO
lVT-77700 GMN N~O IVT-77720 GOR NLO lVT-77145 NLO NLO IVT-77755 OMK NLO IVT-17769 USA NLO
2 1 3 2 3 2 3 2 3
2 3 1 3 2 2
3 2 3
IVT-77821 NLD NLO 2 3 IVT-7782~ NLO NLO 2 2 lYT-77854 NLD NLO 1 3 AC-34e AT~ ATL 2 3 x-71-013 BLV 8LV 2 2
COL.I-136 SPN PER 2 3 eLe CL9 2
C.L.0037 SPN SPN 1 2 C.L.OOS3 PTC PTC 2 3 C.L.Ol.4K PTC PTC 2 2
C.L.0145G pTC PTC 2 .] MEX MEX RS 1 USA USA 2 2
FRC-OQ26 GOR FRC 1 1 FRC-OQ33 FRC 2 3
FRC-004t FRC FRC-OI09 FRC FRC-0144 USA FAC FRC-OI70 R~N FRC FRC-0193 FRC
FRC-0229 F~C FRC FRC-0237 NLO FRC FRC-0292 HLO FR' FRC-0394 SSU FQC FRC-0466 USA FRC
1 2 1 2 2 2 1 2 1 2
1 3 2 2 2 2
2 2 3
FRC-0490 rRC-0491 F"RC-0513 rRe-OS33 PRC-0545
USA FRC 2 uSA FRe 1 1 FPC FAC CF 2 3
FRC CF 1 2 FRC 1 3
1
1
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.------.--------------------------.---------.---.---+--+-+-+--+--,-------+--------------------------+---------+---+---.--+-+-+--+--+
Gl480Q
Gl4b02 Gl4S09
Gl4812 G14619
C I 8 .15 \IIE~ITA
~éTlS T1GRE ROUGE ET 8LANC ~uLrIMA.
FRC-0587 FRC-0590 FRc-0629
FR('-06..:i9
FRC-0669
LflN FRC LeN F Re.
FRC
FRC HK MR GMN f Re
2 2 2 2 2 2 3 3 2 2
1
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G153\Q elS320 (;15323
G15325 el ':-335
G148Z1
G14830 G14~31
G14l:i33 G14B34
'4QIR A EClJSSER o. CHEVILLY FRC-069e. 'ir-oOUBRtCK f'RC-Q897
FRC FRC Mi< 2 L~N FRC 2 1
1 , Gl5335 (;.1 '533 lA G1534Q e15345 Gl 5347
SUPI=.A'METt~
Ti:."IOf"R:ETTr TE~()ERW ... I TE
GI4615 UST~FM ~ARITIA
G14b3b WAIc: 819
G14~~8 YELL~W ~ENE~T
G14ti::'!9 "y G14840 I~A~ E~FAHA.~ lQ
G14d41 I~AN ~OODAROJ 20 G14B42 I~AN GHOChAN 21 G14844 1~~N KARAJ NO.I0 50 Gl4845 IqA~ MINI~TRY NO.136S 51 G14962 JALISCO 42-3
G15004 G15091.A. G15112
'::>15168 G\5228 ~URNL~V 71'óll
GI52:JO il4ALL~lRQuINA
G15234 C.15236
G15248 JAYt;;NA
G15249 KGRA
GI5256 KCRA
G15257 TENOE~ETTE
C.l 5261 <>1'5269 G15.il7J
(.1':'282-Gl~301
G 1 ::>.303 G 1 '5.31 _3 G1531b
F'RC-0907 FRC-OQ2Q F RC.-09.32
FRC FRC '"'~ 2 2 1 , FRC 2 2 U ... A FRC 2. 2 1
FRC-09bl FRC.-0997
FRC-I0Q5
SGA F Re 2 3 G15.3S1 G 15 35 2 G153"S3 G15355 Gl ~d!'l5A
USA FRC 2 2
F~C OT \ 2 , Fr.:C FRC 1 3
Pf416000 lRN U!;A 1 2 3
PI416001 IR"" USA PI416002 IRN USA
Pl416010 IRN USA Pl416011 lRN U~~ MEx-268 MEX CLB
FAO-54.42 C~R ITL FAO-S4&SI C~R lTL FAO-54.54 CMR ITL F40-S4-.60 C!· .. R ITL
AC 36G ATL uTK
C.L.O~17 SP~I ur~
C.L.0423 SPN UTK C.L.Q429 SPN UTK C.L.0453 SPN UTK C.L.u4S5 SPN UTK
C.L.0487 SPN UT~ C&L.04~8 SPN UTK C.L.0502 ~PN UTK C.L.U5~3 SPN UTK C.L.05~1 SPN UTK
C.L.078dO S~")¡~ UTK L~eO-OlI3 zaA ITL ¿Meo-Olll lUA ITL l~8Q-045a ¿BA lTL 1~80-0492 lBA ¡TL
2 1 2 1 2 1 2
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4 G15361 :3 G15365
4 G1537.30
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2 2 G15369 1 2 to4M G15395
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3 1 3 2 2 2 3 2 j
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G15433 WACHS GOLOPERLE G154.14 ALC.Q
2 2 2 3 ,
1 /GlS.47 BISF 1 GIS4S0 CHELINER
G15451 CLA$SJC 3 IGlS454 OORANEL • G15456 OUPRO
2 2 2
12" 2 2 I 1 2 3 2 2 MM 4 1 2 ..
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ELAkN FUGATA LIBAN B& 5 P¡:¡OBATOR H~AN ESF AHA,N 653
2MBO-OSIO ze~ lTL l~60-0554 lBA lTL l~eO-0572 ZBA ITL lM80-0578 IBA 1TL ¿MSO-0705 LBA ITL
Z~80-0712 ZBA lTL z~eQ-Q724 ISA lTL 2MBO-0746 ZBA lTL ¿MBO-OBBS lBA ITL 2MBo-oa93 zeA lTL
LMBO-0990 LBA ITL ZM6Q-0992 IBA ITL M .. BO-006A "'liH 1 T\.. "'W80-007 MWI ITt.. 1<4 ... 80-007 MWI 1 TL
,",wBO-023 MWI 1 TL "'.80-024 MWI lTL MWBO-O¿S MWI ITL ~~80-042 M_l lTL Mw80-174 MWI 1 TL
MweO-311 MWI ITL
MwBO-326 MWI ITL MwaO-354 I'IlWI ITL
M\lJ80-S80 "".-1 I TL MlI\eO-62b MWl lTL
MW60-703 M~l ITL USA USA
IvT-73192 GFR NLo Ivr-73194 GFR NLO IVT-73188 GDR NLO
FRC-0099 FRC FRC FRC-OI74 F~C FRc FRC-OI90 FRC FRC-0354 FRC FRC-0360 NLO FRC
FRC-0373 FRC-0462 FRC-OS86 FRC-07QO PI416284
F .. C
FRC LeN FAC
FRC IR", USA
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3 3 3
2 2 1 2 2 3 2 2
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---------------------------------------------I olplllBI"IH NO. ¡OENTIF1CActON IREGLOCAL R I R le 'FILIE fA
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G'~S82 Gl5586 G155B7 V15S99 <'15593
G15595 Gl'55Q7 GlSb05 G1560b G15608
Tf.I: 33059 TR 33074 TR 3Ju76
TR 33065 TR 33113
Tri 31,189 TR 3_1198
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